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The adenine nucleotide translocation in mitochondria has previously been established as an exchange between exogenous and endogenous adenine nucleotides across the inner membrane. The specificity and the control of the exchange are examined with the following major results: The adenine nucleotide translocation is relatively specific for exogenous ADP and ATP, AMP being nearly inactive. Among other nucleotides tested, only dADP and dATP exchange with a noticeable activity. In the controlled state ADP exchanges 2–4 times faster than ATP. If simultaneously added, ADP and ATP compete for the exchange, with ADP being about tenfold more active than ATP. The specificity of the exit of adenine nucleotides in the exchange is similar to the specificity of the entrance with the difference that ADP and ATP are released with equal activity in proportion to their intramitochondrial content. AMP is released only after a slow conversion to ADP. Therefore the short time exchange is limited by the endogenous content of ADP plus ATP. The exchange is influenced by the metabolic state of the mitochondria. The ATP exchange is more variable than the ADP exchange. Two effects are elucidated: (a) the influence of the metabolic state on the relative content of AMP which inhibits both the ADP and ATP exchange (b) the coupling of the energy transfer system which inhibits only the ATP exchange. An example for case (a) is the inhibition of the ADP and ATP exchange by arsenate and an example for case (b) is the strong increase of the ATP exchange on uncoupling. The following effects are relevant to the mechanism of the control of the exchange by ATP. The stimulation of the ATP exchange by uncoupler has the same concentration dependence as the uncoupling of oxidative phosphorylation (Km [CCP] = 0.08 μM, where CCP = carbonyl‐cyanide‐phenylhydrazone). Oligomycin does not abolish the uncoupler effect on the ATP exchange. “Endogenous uncoupling” on aging of mitochondria also stimulates the ATP exchange. Valinomycin plus K+ only slightly stimulate the ATP exchange. Anaerobiosis stimulates the ATP exchange to a smaller extent than uncoupling.In competition with ADP the effects of energy transfer on ATP exchange are more strongly revealed. On uncoupling the more than tenfold preference for ADP is fully abolished. It is concluded that basically the exchange for ADP and ATP has equal specificity in forward and reverse reaction. In the controlled state a superimposed force makes the specificity asymmetric and inhibits the entrance of ATP. This control of the ATP exchange is concluded to be based on the anionic character of the adenine nucleotides. Thus the ATP4‐ex–ADP3‐in exchange is inhibited unless the charge difference is compensated for by an uncoupler stimulated H+ movement across the membrane. Furthermore an electric potential gradient appears to be effective in the controlled state which is abolished on uncoupling.

The adenine nucleotide translocation in mitochondria has previously been established as an exchange between exogenous and endogenous adenine nucleotides across the inner membrane. The specificity and the control of the exchange are examined with the following major results: The adenine nucleotide translocation is relatively specific for exogenous ADP and ATP, AMP being nearly inactive. Among other nucleotides tested, only dADP and dATP exchange with a noticeable activity. In the controlled state ADP exchanges 2–4 times faster than ATP. If simultaneously added, ADP and ATP compete for the exchange, with ADP being about tenfold more active than ATP. The specificity of the exit of adenine nucleotides in the exchange is similar to the specificity of the entrance with the difference that ADP and ATP are released with equal activity in proportion to their intramitochondrial content. AMP is released only after a slow conversion to ADP. Therefore the short time exchange is limited by the endogenous content of ADP plus ATP. The exchange is influenced by the metabolic state of the mitochondria. The ATP exchange is more variable than the ADP exchange. Two effects are elucidated: (a) the influence of the metabolic state on the relative content of AMP which inhibits both the ADP and ATP exchange (b) the coupling of the energy transfer system which inhibits only the ATP exchange. An example for case (a) is the inhibition of the ADP and ATP exchange by arsenate and an example for case (b) is the strong increase of the ATP exchange on uncoupling. The following effects are relevant to the mechanism of the control of the exchange by ATP. The stimulation of the ATP exchange by uncoupler has the same concentration dependence as the uncoupling of oxidative phosphorylation (Km [CCP] = 0.08 μM, where CCP = carbonyl‐cyanide‐phenylhydrazone). Oligomycin does not abolish the uncoupler effect on the ATP exchange. “Endogenous uncoupling” on aging of mitochondria also stimulates the ATP exchange. Valinomycin plus K+ only slightly stimulate the ATP exchange. Anaerobiosis stimulates the ATP exchange to a smaller extent than uncoupling.In competition with ADP the effects of energy transfer on ATP exchange are more strongly revealed. On uncoupling the more than tenfold preference for ADP is fully abolished. It is concluded that basically the exchange for ADP and ATP has equal specificity in forward and reverse reaction. In the controlled state a superimposed force makes the specificity asymmetric and inhibits the entrance of ATP. This control of the ATP exchange is concluded to be based on the anionic character of the adenine nucleotides. Thus the ATP4‐ex–ADP3‐in exchange is inhibited unless the charge difference is compensated for by an uncoupler stimulated H+ movement across the membrane. Furthermore an electric potential gradient appears to be effective in the controlled state which is abolished on uncoupling.

Cette thèse est basée sur des simulations numériques régionales et aborde, d’un point de vue énergétique, la variabilité à Méso (O(10-100)km) et Sous-mésoéchelle (O(0.1-10)km) dans la région du Courant des Aiguilles.(i) La variabilité de sous-mésoéchelle à la transition entre les deux branches du Courant des Aiguilles (28◦E-26◦E) est dominée, en l’absence de méandres de mésoéchelle, par des tourbillons cycloniques frontaux formant un ‘vortex street’. La tension ambiante frontogénétique intensifie le cisaillement frontal qui déclenche l’instabilité barotrope menant à la génération de tourbillons de sous-mésoéchelle.(ii) Un budget modal d’Energie Cinétique des Tourbillons est développé pour caractériser les transferts d’énergie entre les différentes structures verticales. Les interactions canalisées par la topographie (3 processus) résultent globalement en une perte d’énergie pour les tourbillons de mésoéchelle plus grande que les processus de dissipation (friction au fond et vent) et une cascade verticale inverse (interactions triadiques) renforce les tourbillons de mésoéchelle dans les zones au large.(iii) Notre budget modal permet de caractériser la région du Courant des Aiguilles comme une source nette d’énergie pour les tourbillons de mésoéchelle en contradiction avec celui estimé à partir de données d’altimétrie. Cette différence vient des données d’altimétrie ne tenant pas compte de la contribution principale de la dynamique aux sources et puits d’énergie des tourbillons de mésoéchelle (partie linéaire de la dynamique agéostrophique du mode barotrope et du 1er mode barocline). This dissertation is based on regional numerical simulations and addresses, from an energetic perspective, the Meso (O(10-100)km) and Submesoscale (O(0.1- 10)km) variability in the Agulhas Current region.(i) Submesoscale variability at the transition between the two Agulhas Current branches (28◦E-26◦E) is dominated, in the absence of mesoscale meanders, by cyclonic frontal eddies forming a ’vortex street’. The frontogenetic background strain intensifies the frontal shear which triggers the barotropic instability leading to submesoscale eddies generation.(ii) A modal Eddy Kinetic Energy budget is derived to characterize the energy transfers between the different vertical structures. Interactions canalized by topography (3 processes) globally result in a larger energy loss for mesoscale eddies than dissipation processes (bottom friction and wind) and an inverse vertical cascade (triadic interactions) reinforces mesoscale eddies in offshore areas.(iii) Our modal budget allows to characterize the Agulhas Current region as a net energy source for mesoscale eddies in contradiction with one inferred from altimetry data. The discrepancies come from altimetry data not accounting for the main contribution of the dynamics to mesoscale eddies energy sources and sinks (ageostrophic linear part of the dynamics of the barotropic and 1st baroclinic modes).

Cette thèse est basée sur des simulations numériques régionales et aborde, d’un point de vue énergétique, la variabilité à Méso (O(10-100)km) et Sous-mésoéchelle (O(0.1-10)km) dans la région du Courant des Aiguilles.(i) La variabilité de sous-mésoéchelle à la transition entre les deux branches du Courant des Aiguilles (28◦E-26◦E) est dominée, en l’absence de méandres de mésoéchelle, par des tourbillons cycloniques frontaux formant un ‘vortex street’. La tension ambiante frontogénétique intensifie le cisaillement frontal qui déclenche l’instabilité barotrope menant à la génération de tourbillons de sous-mésoéchelle.(ii) Un budget modal d’Energie Cinétique des Tourbillons est développé pour caractériser les transferts d’énergie entre les différentes structures verticales. Les interactions canalisées par la topographie (3 processus) résultent globalement en une perte d’énergie pour les tourbillons de mésoéchelle plus grande que les processus de dissipation (friction au fond et vent) et une cascade verticale inverse (interactions triadiques) renforce les tourbillons de mésoéchelle dans les zones au large.(iii) Notre budget modal permet de caractériser la région du Courant des Aiguilles comme une source nette d’énergie pour les tourbillons de mésoéchelle en contradiction avec celui estimé à partir de données d’altimétrie. Cette différence vient des données d’altimétrie ne tenant pas compte de la contribution principale de la dynamique aux sources et puits d’énergie des tourbillons de mésoéchelle (partie linéaire de la dynamique agéostrophique du mode barotrope et du 1er mode barocline). This dissertation is based on regional numerical simulations and addresses, from an energetic perspective, the Meso (O(10-100)km) and Submesoscale (O(0.1- 10)km) variability in the Agulhas Current region.(i) Submesoscale variability at the transition between the two Agulhas Current branches (28◦E-26◦E) is dominated, in the absence of mesoscale meanders, by cyclonic frontal eddies forming a ’vortex street’. The frontogenetic background strain intensifies the frontal shear which triggers the barotropic instability leading to submesoscale eddies generation.(ii) A modal Eddy Kinetic Energy budget is derived to characterize the energy transfers between the different vertical structures. Interactions canalized by topography (3 processes) globally result in a larger energy loss for mesoscale eddies than dissipation processes (bottom friction and wind) and an inverse vertical cascade (triadic interactions) reinforces mesoscale eddies in offshore areas.(iii) Our modal budget allows to characterize the Agulhas Current region as a net energy source for mesoscale eddies in contradiction with one inferred from altimetry data. The discrepancies come from altimetry data not accounting for the main contribution of the dynamics to mesoscale eddies energy sources and sinks (ageostrophic linear part of the dynamics of the barotropic and 1st baroclinic modes).

Currently active suspension system is more applicable than passive for improving the suspension performance, ensuring the stability and passenger safety in modern automotive suspension system. However, high energy consumption is one of the main disadvantages of implementing this system in real applications. In this paper, the energy consumption of electromagnetic actuators used for an active suspension system controlled by proportional-integral-derivative (PID), fuzzy adaptive PID, and neuron adaptive PID are investigated through simulation studies. Based on the energy consumption and the performance analysis, it has found that it is potential to develop a vibration energy recovery system to achieve the energy balance requirement in active suspension systems.

Currently active suspension system is more applicable than passive for improving the suspension performance, ensuring the stability and passenger safety in modern automotive suspension system. However, high energy consumption is one of the main disadvantages of implementing this system in real applications. In this paper, the energy consumption of electromagnetic actuators used for an active suspension system controlled by proportional-integral-derivative (PID), fuzzy adaptive PID, and neuron adaptive PID are investigated through simulation studies. Based on the energy consumption and the performance analysis, it has found that it is potential to develop a vibration energy recovery system to achieve the energy balance requirement in active suspension systems.

• Key message Insurance might be an efficient tool to strengthen adaptation of forest management to climate change. A theoretical model under uncertainty is proposed to highlight the effect, on adaptation decisions, of considering adaptation efforts in forest insurance contracts. Results show that insurance is relevant to increase adaptation efforts under some realistic conditions on forest owner’s uncertainty and risk preferences, and on the observability or not of adaptation efforts. • Context One of the challenges of forest adaptation to climate change is to encourage private forest owners to implement adaptation strategies. • Aims We suggest the analysis of forest insurance contracts against natural hazards as a vector to promote the implementation of adaptation efforts by private forest owners. • Methods We propose a theoretical model of insurance economics under risk and under uncertainty. • Results Our results indicate that when climate change makes the probability of the occurrence of the natural event uncertain, then it may be relevant to include adaptation efforts in the insurance contract, leading to an increase in the adaptation efforts of risk-averse and uncertainty-averse forest owners. In addition, we show that the relevance of insurance as a vector to promote adaptation efforts is greater when the forest owner’s effort is unobservable by the insurer as compared to a situation of perfectly observable effort. • Conclusion Under some realistic assumptions, the forest insurance contract seems to be a relevant tool to encourage forest owners to adapt to climate change.